College of Horticulture Science and Engineering, Shandong Agricultural University, Tai'an, 271018, PR China. College of Horticulture Science and Engineering, Shandong Agricultural University, Tai'an, 271018, PR China; State Key Laboratory of Crop Biology, Tai'an, 271018, PR China; Scientific Observing and Experimental Station of Environment Controlled Agricultural Engineering in Huang-Huai-Hai Region, Ministry of Agriculture, Tai'an, 271018, PR China. Electronic address: gslqm@sdau.edu.cn. State Key Laboratory of Crop Biology, Tai'an, 271018, PR China. Electronic address: lbroom@163.com. College of Horticulture Science and Engineering, Shandong Agricultural University, Tai'an, 271018, PR China; State Key Laboratory of Crop Biology, Tai'an, 271018, PR China. College of Horticulture Science and Engineering, Shandong Agricultural University, Tai'an, 271018, PR China; State Key Laboratory of Crop Biology, Tai'an, 271018, PR China; Scientific Observing and Experimental Station of Environment Controlled Agricultural Engineering in Huang-Huai-Hai Region, Ministry of Agriculture, Tai'an, 271018, PR China.